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Verity L. Mathis, Mark S. Hafner, David J. Here we reevaluate relationships within this species, which, based on previous studies, is composed of 3 genetically divergent clades. Reanalysis of published allozyme data shows a pattern of isolation by distance consistent with the hypothesis of gene flow among the 3 T. Species tree analyses of these clades reveal 4 divergent lineages 2 within the TMVB clade , which we recognize herein at the subspecies level. Cranial morphometries show a moderate level of morphological separation among these clades in multivariate space.
The present center of diversity of the T. A synonymy of T. El centro actual de diversidad del grupo de especies de T. Una sinonimia para T. A subspecies can be viewed as either a unit of classification to describe geographical variation within a species or as an evolving lineage potentially on a trajectory toward speciation Lidicker Although these definitions are not mutually exclusive, systematists rarely publish explicit statements about their rationale for describing and naming subspecies.
For many taxa, morphological variation may be indicative of incipient speciation Zimmerman et al. In others, morphological differences that define subspecies may not coincide with molecular breaks within the species Conroy and Cook ; Conroy and Neuwald The complexities of mosaic evolution coupled with divergent views on the importance of variation at different levels of biological organization make it unlikely that researchers will agree on a single, universal subspecies concept.
This makes it imperative that taxonomic studies at the intraspecific level include a clear statement of the author's concept of subspecies to provide a context in which to evaluate the robustness and defensibility of the author's taxonomic decisions. Taxonomy of the smooth-toothed pocket gophers, Thomomys , has long been fraught with conflict over species and subspecies boundaries.
